As reported by the Entomology Society of America, the Formosan subterranean termite (Coptotermes formosanus) is an interesting model for studying the evolution of termites and complex behaviors in eusocial insects.
ESA cited a study published in January in Environmental Entomology that investigated the social behaviors of workers in young and old termite colonies and showed that, as the termite colony matures, the task division according to the age of the workers becomes more complex.
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Source: Entomology Today